Muscular Tissues
Contractility and motility (movement) are fundamental properties of protoplasm. That is why, all cells possess potential motility. Contraction for motility in the cells results essentially from the interaction of two contractile proteins, actin and myosin. These proteins enter into the composition of microfilaments of cellular cytoskeleton. During evolution, organism achieved enhancement in motility by various means. For example, unicellular organisms and cells of lower metazoans (mesozoa and parazoa) acquired the ability to form pseudopodia, flagella, or cilia as locomotory organelles. Then, in eumetazoans began the evolution of specialized contractile cells having much more of actin and myosin proteins. In the lowest eumetazoans (cnidarians), basal ends of epithelial cells are drawn out into elongated contractile processes. That is why, these cells are called epitheliomuscular or myoepithelial cells. In the eumetazoans higher than cnidarians, independent contractile cells, called muscle cells (myocytes), occur and form Muscular Tissues (muscles). These tissues are obviously responsible for movements of organs and locomotion of the body in response to stimuli. These develop from embryonic mesoderm except for those of the iris and ciliary body of eyes, which are ectodermal in origin. About 40% to 50% of our body mass is of muscles. The muscle cells are always elongated, slender and spindle-shaped, fibre-like cells, These are, therefore called muscle fibres. These possess large numbers of myofibrils formed of actin and myosin. The myofibrils are obviously stuffed in a small amount of cytoplasm due to thin, fibre-like shape of muscle cells. Muscle cells are highly contractile (contracting to 1/3 or 1/2 the resting length). Muscle cells lose capacity to divide, multiply and regenerate to a great extent. Study of muscle is called myology. Types of muscle are following -
(i) Striated or striped muscles : Most muscles of body are striated. These generally bring about voluntary movements under conscious control of brain and, hence, called voluntary muscles. Most of these are inserted at both ends upon bones in different parts of the body depend upon these muscles. Hence, these are also called skeletal muscles. Movements of limbs and the body solely depend upon these muscles. Hence these are also called somatic muscles. These are also called phasic type of muscles, because contraction in these is rapid, but brief and fatigue occurs quickly.
(a) Structure of striated muscles:
Each striated muscle consists of numerous muscle fibres segregated into several small and parallel bundles, called Fasciculi. Fibres of each fascicule are bound together by a connective tissue sheath, called endomycium. All fasciculi of a muscle are bound together by a connective tissue termed perimycium which also forms a sheath around each fascicule. Similarly, the whole muscle itself is covered by a connective tissue sheath, called epimycium. The latter extends as a tendons at each end of the muscle to insert it on to bones. Endomycium,
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