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First Law of Thermodynamics   First law of thermodynamics was proposed by Helmholtz and Robert Mayer. This law is also known as law of conservation of energy. It states that, “Energy can neither be created nor destroyed although it can be converted from one form into another.” (1) Justification for the law : The first law of thermodynamics has no theoretical proof. This law is based on human experience and has not yet been violated. The following observations justify the validity of this law (i) The total energy of an isolated system remains constant although it can undergo a change from one form to another. (ii) It is not possible to construct a perpetual machine which can do work without the expenditure of energy, If the law were not true, it would have been possible to construct such a machine.           (iii) James Joule (1850) conducted a large number of experiments regarding the conversion of work into heat energy. He concluded that for every 4.183 joule of work done on the system, one calorie of heat is produced. He also pointed out that the same amount of work done always produces same amount of heat irrespective of how the work is done.    (iv) Energy is conserved in chemical reactions also. For example, the electrical energy equivalent to 286.4 kJ mol-1 of energy is consumed when one mole of water decomposes into gaseous hydrogen and oxygen. On the other hand, the same amount of energy in the form of heat is liberated when one mole of liquid water is produced from gases hydrogen and oxygen. \[{{H}_{2}}O(l)+286.4\,kJ\xrightarrow{{}}{{H}_{2}}(g)+\frac{1}{2}{{O}_{2}}(g)\];\[{{H}_{2}}(g)+\frac{1}{2}{{O}_{2}}(g)\xrightarrow{{}}{{H}_{2}}O(l)+286.4\,kJ\] These examples justify that energy is always conserved though it may change its form. (2) Mathematical expression for the law : The internal energy of a system can be changed in two ways (i) By allowing heat to flow into the system or out of the system. (ii) By doing work on the system or by the system. Let us consider a system whose internal energy is \[{{E}_{1}}\]. Now, if the system absorbs q amount of heat, then the internal energy of the system increases and becomes \[{{E}_{1}}+q\]. If work \[(w)\]is done on the system, then its internal energy further increases and becomes \[{{E}_{2}}\]. Thus,                                                 \[{{E}_{2}}={{E}_{1}}+q+w\] or \[{{E}_{2}}-{{E}_{1}}=q+w\] or \[\Delta E\,=q+w\] i.e. \[(Change\,in\,internal\,energy)\]=\[(Heat\,added\,to\,the\,system)+(Work\,done\,on\,the\,system)\] If a system does work (w) on the surroundings, its internal energy decreases. In this case, work is taken as negative (–w). Now, q is the amount of heat added to the system and w is the work done by the system, then change in internal energy becomes, \[\Delta E=q+(-w)=q-w\] i.e. \[(Change\,in\,internal\,energy)\]=\[(Heat\,added\,to\,the\,system)-(Work\,done\,by\,the\,system)\] The relationship between internal energy, work and heat is a mathematical statement of first law of thermodynamics. (3) Some useful conclusions drawn from the law :  \[\Delta E=q+w\] (i) When a system undergoes a change \[\Delta E=0\], i.e., there is no increase or decrease in the internal energy of the system, the first law of thermodynamics reduces to                                                 \[0=q+w\]  or  \[q=-w\]             (heat more...

  Asexual Reproduction   Asexual reproduction. The methods of reproduction which do not involve meiosis and fertilization are known as apomixis or asexual reproduction. Only mitotic divisions are involved in these methods, resulting into the formation of offsprings which are genetically similar to the parent plant.   Asexual reproduction is of following two types: (1) Agamospermy: Agamospermy is a kind of plant apomixis in which the embryos and seeds are formed by asexual reproductive methods without involving meiotic gametogenesis and sexual fusion of gametes. It occurs widely in ferns and the flowering plants. There are three different types of agamospermy:         (i) Adventive embryony : Formation of embryo directly from the diploid sporophytic cells (nucellus or integument) of ovule is called adventive embryony. Such embryos are formed without involving meiosis and sexual fusion, e.g., Citrus, Opuntia, etc. In Citrus, a seed may possess upto 40 embryos (one normal and rest adventive). (ii) Diplospory : In this case, the archesporium differentiates but megaspore mother cell directly gives rise to an unreduced (i.e., without meiosis) embryo sac. It may produce two types of embryos : (a) Diploid parthenogenesis : Embryo develops from unfertilized diploid egg. (b) Diploid apogamy : Embryo develops from any diploid cell of embryo sac except egg. (iii) Apospory : It is the formation of complete embryo sac from the sporophytic cell without meiosis so that the gametophyte remains diploid. Apospory may be of two types : (a) Somatic apospory : Embryo sac is formed from somatic cell. (b) Generative apospory : Embryo sac is formed from archesporium without meiosis.   (2) Vegetative propagation: Regeneration or Formation of a new individual from any vegetative part of the body is called vegetative reproduction or vegetative propagation. The lower plants reproduce vegetatively through budding, fission, fragmentation, gemmae, resting buds, spores, etc. Among flowering plants, every part of the body such as roots, stem, leaves and buds take part in vegetative propagation. It is very common mode of reproduction and it may be natural vegetative propagation or artificial vegetative propagation. (i) Natural methods of vegetative propagation : In natural vegetative propagation, a portion gets deattached from the body of mother plant and it grows into a new individual plant under suitable conditions. Different plant parts are variously modified for vegetative propagation. Some of these are given below. (a) Vegetative propagation by stems: The modified stems like bulbs, runners, rhizomes, corms, tubers, offsets, etc., help the plant to multiply under favourable conditions.
  • Bulb: It is a modified shoot that has a very short stem and apical and axillary buds. Some of these grow to form shoots. g. Onion, Tulip, Lilies, Garlic, etc.
  • Runners: These are creeping modified stems which produce adventitious roots at nodes. Each node gives rise to aerial shoot which becomes a new plant g. Doob grass (Cynodon), Wood sorrel (Oxalis), Indian pennywort (Centella), etc.
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  Microsporogenesis   Microsporogenesis The process of the formation and differentiation of microspores (pollen grains) from microspore mother cells (MMC) by reductional division is called microsporogenesis. Microsporogenesis is well studied under following heads:       (1) Structure of anther: The fertile portion of stamens is called anther. Each anther is usually made up of two lobes connected by a connective. In turn each anther lobe contains two pollen chambers placed longitudinally. Each pollen chamber represents a microsporangium and is filled with a large number of pollen grains or microspores. A typical anther consist of four microsporangia (tetrasporangiate) and such anthers is called dithecous e.g. mostly plants. In members of Malvaceae anthers are reniform or kidney shaped and consist of two microspoangia (bisporangiate), such anthers is called monothecous. In the smallest parasitic angiosperm, Arceuthobium minutissimum, anthers consist of only one microsporangium (monosporangiate). The pollen sacs are surrounded by following 4 layers : (i) Epidermis : This is the outermost single layered and protective. In Arceuthobium, cells of epidermis develops a fibrous thickening and the epidermis is designated as exothecium. (ii) Endothecium : Inner to epidermis, there is a single layer of radially elongated cells. Cells of endothecium develop fibrous thickening (made up of cellulose with a little pectin and lignin) which help in the dehiscence of anther. In between these cells, a few cells without thickening are also present. These thick walled cells collectively form the stomium. (iii) Middle layer : Three to four layers of thin walled cells situated just below the endothecium are known as middle layers. Cells of this layer are ephemeral and degenerate to provide nourishment to growing microspore mother cells. (iv) Tapetum : This is the innermost layer of the wall. The cells are multinucleate (undergo endopolyploidy) and polyploid. Tapetal cells are nutritive. In these cells the Ubisch bodies are present which help in the ornamentation of microspore walls. A compound sporopollenin is secreted in the exine of microspore wall. According to Periasamy and Swamy (1966), developmentally the tapetum has dual nature. The tapetum is of two types : (a) Amoeboid or Periplasmodial tapetum : In young condition cell wall of tapetal cells breaks, so protoplast of these cells become free between microspore mother cell and form mass of tapetal periplasmodium. e.g. Alisma, Typha, Tradescantia. (b) Secretory or Glandular tapetum : This is the most common type of tapetum which remains insitu as such throughout. The tapetal cells secretes nourishment that passes into sporogenous cells. This tapetum attains its maximum development at the stage of pollen tetrads and then degenerates.   (2) Development of anther and formation of microspores (Pollen grains): The young anther consists of homogenous mass of paranchymatous cells surrounded by epidermis. It soon becomes four lobed. In each of the four lobes, some of the hypodermal cells begin to act as archesporial initials. Each archesporial initial divides into an outer more...

  Megasporogenesis   Megasporogenesis The process of formation of megaspore from megaspore mother cell by meiotic division is known as megasporogenesis. This process takes place in ovule. Megasporogenesis can be studied under following heads:                                                                                                                                                                                                                                             (1) Structure of ovule: Ovule is considered to be an integumented megasporangium. The ovule consists of the stalk and the body. The stalk is called funicle. One end of the funicle is attached to placenta and the other end to the body of the ovule. The point of attachment of funicle with the body is called hilum. Sometimes funicle gets fused with the body of the ovule one side and forms a ridge known as raphe. The body of the ovule shows two ends: the basal end, often called the chalazal end and the upper end is called micropylar end. The main body of the ovule is covered with one or two envelopes called integuments. These leave an opening at the top of the ovule called micropyle. The integuments enclose a large parenchymatous tissue known as nucellus. The residual part of nucellus in the mature seed is called perisperm. In the centre of the nucellus is situated a female gametophyte known as embryo sac. Following are the conditions seen in ovule in relation to integuments: (i) Unitegmic : Ovule with a single integument, e.g., sympetalous or gamopetalous dicotyledons. (ii) Bitegmic: Ovule with two integuments as in polypetalous (Archichlamydeae) dicotyledons and monocotyledons. (iii) Aril : This is a collar-like outgrowth from the base of the ovule and forms third integument. Aril is found in litchi, nutmeg, etc. (iv) Caruncle : It is formed as an outgrowth of the outer integument in the micropylar region. Caruncle is common in the ovules of Euphorbiaceae. e.g., Castor (Ricinus). (v) Ategmic : In some parasites like Loranthus, Viscum, Santalum etc., there is no integument. Such an ovule is called ategmic. (2) Kinds of ovules: Depending upon the shape and orientation, the ovules of angiosperms are classified into following types : (i) Orthotropous or Atropus : The micropyle, chalaza and funicle are in straight line. This is most primitive type of ovules. e.g. Betel, Piper, Polygonum. (ii) Anatropous : The body of the ovule is completely inverted (turn at 180o angle ) so that micropyle and hilum come to lie very close to each other. e.g. 82% of angiosperm families. (iii) Hemianatropous : Ovule turns at 90o angle upon the funicle or body of ovule is at right angle to the funicle e.g. Ranunculus. (iv) Campylotropous : Ovule is circled more or less at right angle to funicle. Micropylar end is bent down slightly. e.g. in members of Leguminaceae and Cruciferae.     more...

  Pollination   Pollination The process of transfer of pollen grains, from an anther to the stigma of the same flower or of different flower. It is of two types: (1) Self pollination                       (2) Cross pollination     (1) Self-pollination: This process involves the transfer of pollen grains from the anthers to the stigma of the same flower or of another flower borne by the same plant. It is of two types: (i) Autogamy : It is a kind of pollination in which the pollen from the anthers of a flower are transferred to the stigma of the same flower. (ii) Geitonogamy : It is an kind of pollination in which the pollen from the anthers of one flower are transferred to the stigma of another flower borne on the same plant. It usually occurs in plants which show monoecious condition (unisexual, male and female flowers are borne on the same plant). Geitonogamy involves two flowers but these belong to the same parent plant. Merits
  • Pollen grains are not wasted.
  • The purity of the generation is maintained.
Demerits
  • New and healthier varieties are not formed.
  • It results in weaker progeny, producing weaker seeds and plants.
Contrivances for self-pollination: The major contrivances or adaptations which favours self-pollination are :  (a) Bisexuality : Flowers should be bisexual or hermophrodite. (b) Homogamy : Anthers and stigma of the bisexual flowers of some plants mature at the same time. They are brought close to each other by growth, bending or folding to ensure self-pollination. This condition is called homogamy. e.g., Mirabilis (Four O, clock), Catharanthus (= Vinca), Potato, Sunflower, Wheat, Rice, etc. (c) Cleistogamy : Some plants never open to ensure complete self-pollination. This condition is called cleistogamy, e.g., Commelina bengalensis, Oxalis, Viola, etc. The cleistogamous flowers are bisexual small, inconspicious, colourless and do not secrete nectar.   (2) Cross pollination: Cross pollination involves the transfer of pollen grains from the flower of one plant to the stigma of the flower of another plant. It is also called xenogamy.   Merits  
  • Seeds are more and viable.
  • Progenies are healthier.
  • Adaptability is better.
  • New varieties can be produced.
  Demerits
  • The process is not definite because plants depend on agencies.
  • Large amount of pollen grains are wasted.
Contrivances for cross pollination: Nature favours cross pollination. All unisexual flowers and a large number of bisexual flowers are naturally cross pollinated. The main contrivances ensuring cross pollination are as follows: (i) Diclincy or Unisexuality : In unisexual flowers stamens and carpels are found in different flowers. Unisexuality can be of two types :
  • Monoecious plant : When male and female flowers are borne on the same plant. g., Maize, Cucurbits, Castor.
  • Dioecious plant : When male and female flowers are borne on different plants. g., more...

  Fertilization   Fertilization The fusion of two dissimilar sexual reproductive units (gametes) is called fertilization. This process was discovered by Strasburger (1884). (1) Germination of pollen grain on stigma and growth of pollen tube: Pollen grains reach the receptive stigma of the carpel by the act of pollination. Pollen grains, after getting attached to the stigma, absorb water and swell. Subsequent to mutual recognition and acceptance of pollen grains, the pollen grain germinates (in vivo) to produce a pollen tube which grows into stigma towards the ovarian cavity. G.B. Amici (1824) discovered the pollen tube in Portulaca oleracea. Generally, only one pollen tube is produced by a pollen grain (monosiphonous). But in some plants like members of Cucurbitaceae produce many pollen tubes (polysiphonous). The pollen tube contains a vegetative nucleus or tube nucleus and two male gametes. Later, the vegetative cell degenerates. The pollen tube now reaches the ovule after passing through the style.         (2) Entry of pollen tube into ovule: After reaching ovary, the pollen tube enters the ovule. Pollen tube may enter the ovule by any one of the following routes:   (i) Porogamy : When the pollen tube enters the ovule through micropyle, it is called porogamy. It is the most common type. e.g. Lily.   (ii) Chalazogamy : The entry of pollen tube into the ovule from chalazal region is known as chalazogamy. Chalazogamy is less common. e.g. Casuarina, Juglans, Betula, etc. It was first observed by Treub (1981) in Casuarina.   (iii) Mesogamy : The pollen tube enters the ovule through its middle part i.e. through integument (e.g. Cucurbita, Populus) or through funicle (e.g. Pistacia).   (3) Entry of pollen tube into embryo sac: The pollen tube enters the embryo sac only from the micropylar end irrespective of its mode of entry into the ovule. The pollen tube either passes between a synergid and the egg cell or enters into one of the synergids through filiform apparatus. The synergids direct the growth of pollen tube by secreting some chemical substances (chemotropic secretion). The tip of pollen tube enters into one synergid. The penetrated synergid starts degenerating. After penetration, the tip of pollen tube enlarge and ruptures releasing most of its contents including the two male gametes and the vegetative nucleus into the synergid.   (4) Double fertilization: The nuclei of both the male gametes are released in the embryo sac. One male gamete fuses with the egg to form the diploid zygote. The process is called syngamy or generative fertilization. This syngamy was discovered by Strasburger (1884). The diploid zygote finally develops into embryo. The other male gamete fuses with the two polar nuclei (or secondary nucleus) to form the triploid primary endosperm nucleus. The process is called triple fusion or vegetative fertilization. These two acts of fertilizations constitute the process of double fertilization. The process was discovered more...

  Embryo   Embryo (1) Development of embryo (Embryogeny): The zygote after a period of rest develops into embryo. The process of development of mature embryo from diploid zygote is called embryogenesis. (i) In dicotyledons : The normal type of dicot embryo development has been studied in Shephered purse  (Capsella bursapastoris) family Cruciferae. This is called as crucifer or onagrad type of embryo development. This development of embryo is endoscopic i.e. apex is downward or towards inside. The first division of zygote is transverse which produces a basal cell (cb) towards the micropyle and a terminal cell (ca) towards chalaza. The basal cell divides by transverse division and the terminal cell by a longitudinal division, so 4 celled T-shaped proembryo is produced. The two basal cells divide by transverse division and form 6-10 celled suspensor. The upper most cell of the suspensor is vasicular cell and lowest cell is called hypophysis which forms radicle and root cap.         The two apical cells first divide by longitudinal division (at right angle to first one) and then by transverse and periclinal division. So sixteen celled globular embryo is produced. Due to differentiation of cotyledons globular embryo becomes heart shaped. Mature embryo in dicots consists of two lateral cotyledons, terminal plumule or stem tip and radicle or root tip.   (ii) In monocotyledons: The normal type of monocot embryo development has been studied in Sagittaria sagittaefolia. The early development of dicot and monocot embryos is similar upto globular stage. Later on differentiation starts. Suspensor is single celled and vascular. There is only one terminal cotyledon called scutellum (shield shaped). In grasses the second cotyledon is reduced called epiblast.           The basal cell (cb) divides by a transverse wall into two cells ­– ci and m. The cell ci divides once again to form n and n’ cells. Of these n’ is the outermost which develops into suspensor. The cell n forms parts of root cap the cell m contributes to the remaining part of root cap and a part of the radicle. The terminal cell (ca) divides by two vertical walls, at right angles to one another. This results in the formation of a quadrant (q). Cells of the quadrant divide periclinally differentiating into the peripheral cells and the inner group of cells. The repeated divisions in both peripheral and central group of cells results in the formation of two regions –l and l’. Region l produces the lower part of cotyledon while upper part of cotyledon, hypocotyl and plumule are formed by l’ region.   (2) Polyembryony: Occurrence of more than two embryo in the seed is known as polyembryony. It was discovered by A.V. Leeuwenhock (1719) in Citrus. It may be :   (i) Cleavage polyembryony more...

  Parthenocarpy       Parthenocarpy The formation of fruits without fertilization is called parthenocarpy. Such fruits are either seedless or non-viable seeds. Parthenocarpy is of two types:   (1) Natural parthenocarpy: When seedless fruits are produced without any special treatment from the ovaries in the absence of pollination and fertilization, the phenomenon is called natural parthenocarpy. e.g., Grapes, Banana, Pineapple and Noval oranges.   (2) Induced parthenocarpy: When seedless fruits are produced by spraying the flowers with either water extract of pollen grains or growth promoting hormones such as Indole acetic acid (IAA), Naphthalene acetic acid (NAA), Gibberellic acid (GA), etc. the phenomenon is called induced parthenocarpy. e.g., Tomato, Black berry, Fig, Lemon, Apple, Orange, Pear. etc.

  Morphology of Frog     Introduction Frogs are carnivorous tailless amphibians which are widely found in India. A diverse variety of frogs can be found all over the world; among them, the Indian frogs are called Rana tigirna. They are vertebrates, coming under the class Amphibia (phylum Chordata). Frogs are cold-blooded animals (poikilotherms) whose body temperature varies according to their environment. Hence, they need to protect themselves from extreme heat and cold for maintaining optimum body temperature. Thus, they follow aestivation and hibernation during summer and winter seasons. Another characteristic feature of frogs is that they are camouflage i.e., they can change their skin color according to their surroundings. Related image Morphology of Frogs   Though larvae have tails, adult frogs are tailless. An adult frog has a stout body which is differentiated into head and trunk. Other external features are a pair of nostrils, protruding eyes, a membranous tympanum (ear), slippery/warty moist skin and webbed limbs.   Frogs   Frogs generally have a slippery moist and highly permeable skin through which they absorb water and respire. Thus, the moist skin acts as a respiratory organ in frogs. Also, the skin is glandular in nature which produces mucus and toxic substances to warn them of their predators. The color of the skin can vary from brown and green to vivid colors as per secretions.   The locomotion of frogs takes place with the help of their forelimbs and hind limbs. Frogs are unisexual i.e., they show sexual dimorphism. A male frog is distinguished from a female frog by the presence of vocal sacs and a copulatory pad on forelimbs. A female frog lacks these body features.   Anatomy   The body plan of frogs consists of well-developed structures which help them in their physiological activities. The body cavity accommodates all the organ systems such as digestive, respiratory, circulatory, excretory, nervous and reproductive systems whose functions are almost similar to human body systems.   Digestive system:   The alimentary canal together with the accessory organs makes up the digestive system of the frog. Since frogs are carnivorous they have short intestine. The alimentary canal begins at the mouth (buccal or oral cavity), passes through the pharynx, esophagus or food pipe, stomach, small intestines, large intestines, rectum and finally ending at the cloaca. The food particles get digested gradually as they travel through various compartments of the alimentary canal. more...

  Smoking   Smoking of tobacco dried and cured leaves of plant ‘Nicotiana tobacum’ and N. rustica in the form of cigars, cigarettes, bides etc. is very toxic to the body. Smoke of tobacco contains about 300 compounds. The main compounds are nicotine, CO, HCN, polycyclic aromatic hydrocarbons, certain other stimulating products etc. Nicotine in the blood stimulates the nervous system, relax the muscles, release adrenalin hormone and increase the rate of heart beats. In pregnant ladies growth of fetus decreases and loss of weight may takes place.   (i) Tobacco: The tobacco was first smoked by Red Indians in America. It then spread to European countries in the early 1600’s, and today a large part of the world population smoke tobacco, while some others chew it.   (ii) Effect of Nicotine: Smoking was reported to produce a feeling of tranquility (calmness) and in some cases made people alert and active. Since in its early days the use of tobacco was socially accepted and no harmful effects were obvious, the addiction became widespread. But scientific research indicates that use of tobacco is harmful. Nicotine is the major stimulatory component of tobacco products including cigarettes. It is highly poisonous. The amount present in one cigar can be fatal, if it is injected intravenously into a person. When smoked, about 10 percent of the smoke is inhaled. Nicotine has a number of effects on the human body. It stimulates passage of nerve impulses, causes muscles to relax and causes the release of adrenaline, increasing both blood pressure and heart beat rate. The increased blood pressure caused by smoking leads to increased risk of heart diseases. In pregnant women nicotine causes retardation of the growth of the foetus.   Other Harmful Components of Tobacco Smoke: Besides the nicotine the tobacco smoke contains carbon monoxide, polycyclic aromatic hydrocarbons and tar.   (iii) Deseases Caused by Smoking: Smoking causes the following diseases. (a) Cancer: Benzpyrene present in tobacco smoke is carcinogenic. About 95% victims of lung cancer are due to smoking. Reverse smoking causes oral cancer. In reverse smoking the burning end of the cigar is kept in the mouth. Reverse smoking is common in the villages of Andhra Pradesh. Bidi smoking causes cancer of tongue, pharynx (throat), larynx, tonsils and oesophagus. Lip cancer is caused by cigar and pipes. Tobacco chewing leads to oral cancer. Tobacco smoking mutates and inactivates P 53 gene which checks cancer growth. Carcinogenic agents are X-ray radiation, U.V. radiation, Nicotine etc. (b) Cardio-vascular Diseases: Tobacco smoking causes increase of adrenalin secretion which increases blood pressure, heart beat rate by constricting the arteries. High blood pressure increases the chances of heart diseases. Nicotine damages the bicuspid valve (mitral valve) of the heart. Nicotine acts as a stimulant because it minimize the effect of acetylcholine. (c) Emphysema: Tobacco smoke may break down the walls of alveoli of the lungs, decreasing the surface area for gas exchange, causing more...


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