Water as an important ecological factor was first recognized by Warming. On the basis of water requirement plants are divided in to 3 major types :
(1) Hydrophytes : They live in abundance of water. They require abundance of water to complete their life cycle. They are of following types :
(i) Rooted submerged : The hydrophytes, which are rooted in mud at bottom and remain under water. e.g., Hydrilla, Vallisnaria.
(ii) Submerged floating : They are not rooted in the soil but completely submerged and floating e.g., Ceratophyllum, Utricularia.
(iii) Rooted with floating leaves : They are rooted in the soil but the leaves are floating on the surface of water e.g., Nelumbo, Trapa, Victoria.
(iv) Free floating : They are not rooted in soil and mud. These hydrophytes float freely on the surface of water e.g., Wolfia (Smallest angiosperm), Lemna, Spirodella, Pistia, Azolla, Salvinia.
(v) Rooted emergent : Roots are in soil shoots or leaves are partly outside and partly inside the water. Plants show heterophilly (Amphibious plants) e.g., Typha, Ranunculus, Sagittaria, Cyperus.
Morphological adaptations
(i) Roots of hydrophytes are poorly developed or completely absent in Wolfia, Ceratophyllum etc. Root hair absent but root pockets may be present e.g., Pistia, Eichornia, Trapa.
(ii) Stem is reduced in free floating plants e.g., Pistia, narrow and slender in submerged plants e.g., Hydrilla, Ceratophyllum and well developed in amphibious plants e.g., Typha.
(iii) Petioles become long, swollen and spongy for floating.
(iv) Leaves are usually long ribbon like e.g., Potamogeton, or finely divided e.g., Ranunculus or thin and broad e.g., Nelumbo, Victoria.
(v) In some hydrophytes leaves of different forms are produced by same plant. Aerial leaves are not dissected but submerged leaves dissected (Heterophilly) e.g., Ranunculuc, Limnophila.
Anatomical adaptations
(i) Cuticle absent or poorly developed.
(ii) Stomata are absent in submerged plants. Floating hydrophytes have stomata on upper surface e.g., Lotus (epistomatic).
(iii) Air spaces are extensively developed in root, stem and leaves. Well developed aerenchyma helps in buoyancy and gaseous exchange.
(iv) Leaves have spongy tissues and palisade is poorly developed. As light difuses from all palisade and spongy tissue. Epidermal cells contain chloroplasts for maximum capturing of difused light.
(v) Mechanical tissues like sclerenchyma (lignified tissues) and collenchyma are poorly developed or absent.
(vi) Vascular tissues are poorly developed.
Physiological adaptation
(i) Water and mineral nutrients are absorbed through general body surface.
(ii) Osmotic concentration or osmotic potential of cells is equal to or is slightly higher than external water.
(2) Xerophytes : They are adapted to grow in dry habitats. On the basis of pattern of life cycle, xerophytes are of three types :
(i) Ephemerals : They complete their life cycle in a very short period, evade dry season by disappearing, leaving their seeds. They are referred as drought escapers or drought evaderes e.g., Cassia toria, Argemone maxicana, Solanum xanthocarpum.
(ii) Succulents (Fleshy xerophytes) : They absorb large quantities of water during rainy season and store water in different body parts. They are common in deserts
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